On a boat, the two cross a river, and after climbing a vine, reach Truffle Towers. Mario decides to go stop Waluigi and Toad decides to come with him to Truffle Towers. Toad warns Mario (or Luigi) that someone has stolen the Music Keys and explains the trouble that this causes. Meanwhile, Toad, having seen this, rushes to tell Mario or Luigi, depending on which character the player chose. However, when Waluigi opens the door to the room of the four Music Keys, they all scatter across the Mushroom Kingdom except for one that Waluigi gets to keep. This is trouble as the Music Keys are able to grant any wish. The opening scene starts out with Waluigi breaking into Truffle Towers. The playable characters are Mario and Luigi. The game includes a Story Mode, which the player must clear in order to unlock all of the songs in the game. Toadette confronting the two protagonists.
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